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SUMMARY:Specification of pre-meiotic cells in maize anthers - Virginia Wal
 bot\, Stanford University
DTSTART:20120308T160000Z
DTEND:20120308T170000Z
UID:TALK31564@talks.cam.ac.uk
CONTACT:Jill Harrison
DESCRIPTION:Without a dedicated germ line\, how do plant cells differentia
 te for meiosis?  We are studying the steps in maize anther ontogeny from a
 nther primordium inception until the onset of meiosis.  The first step is 
 production 4 lobes on each anther establishing the locules and overall ant
 her morphology.  Within the locules we have charted the successive specifi
 cation of cell types:  in order of appearance epidermal cells from the L1 
 and in the L2 the archesporial\, then the endothecial and secondary pariet
 al cell types\, and finally the middle layer and tapetal cell types.  Usin
 g confocal reconstruction and analysis of maize mutants\, we have develope
 d a new theory for the specification of the archesporial (pre-meiotic cell
 s) and somatic cell types to replace the classic lineage model.  These mic
 roscopic analyses are complemented by the cloning of key genes controlling
  steps in anther cell differentiation and by microarray and proteomic anal
 yses of whole anthers and individual cell types.   We have also discovered
  physiological conditions that can overcome mutations in cell fate specifi
 cation and propose that a combination of developmental and physiological c
 ues operate in situ during key steps in anther cell fate determination.
LOCATION:Sainsbury Laboratory\, Cambridge University\, Bateman Street
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