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SUMMARY:Building plant bodies from hair - Professor Liam Dolan\, John Inne
 s Centre\, Norwich
DTSTART:20080221T160000Z
DTEND:20080221T170000Z
UID:TALK9399@talks.cam.ac.uk
CONTACT:3901
DESCRIPTION:Root hairs are required for the uptake of minerals and water f
 rom soil and for anchorage during growth and development of the root syste
 m in many plant species. Hairs elongate by a mechanism known as tip growth
 \, where cell expansion is focussed to a restricted region of the cell sur
 face. Similar cell types are found throughout the land plants. For example
  caulonema and rhizoids are tip-growing cells found in the mosses. In the 
 ferns tip growing rhizoid cells are found in both gametophyte and sporophy
 te generations. This indicates that the production of tip-growing cells th
 at interact with the substrate are found throughout the land plants. Furth
 ermore\, their occurrence in the green algae indicates that the mechanism 
 by which they develop may be ancient.\n\nA cascade of transcription factor
 s regulates the development of root hairs in the model angiosperm Arabidop
 sis. Early acting transcription factors control the fate of cells in the r
 oot epidermis - epidermal cells may be hair-bearing or hairless. We have e
 vidence that the early acting genes also control the expression of a suite
  of late acting transcription factors that are required for late stages of
  root hair differentiation when hair outgrowth occurs. One of these genes\
 , RHD6\, is only expressed in the root hair cell where it promotes the tra
 nscription of three other related genes which are also required for root h
 air cell differentiation. Evidence will be presented that defines the regu
 latory interactions between these genes in Arabidopsis and I will demonstr
 ate that RHD6 and RHD6-LIKE genes are part of an ancient mechanism that co
 ntrols the differentiation of tip-growing cells in plants.\n\nhttp://www.j
 ic.ac.uk/staff/liam-dolan/
LOCATION:Department of Plant Sciences\, Large Lecture Theatre
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